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If I were to replace most of my upper gearbox, would it be a better idea to go with the ICS MA-54/56 complete uppers, replace the piston with a polycarb, then downgrade the spring (say M100/M110) and leave it at that? Would the stock gears be fine handling that sort of setup?
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You mean the upper mechbox, yes? We're talking about an ICS M4. One of the biggest features of the ICS M4 is the split mechbox and the option for you to swap out your previous upper mechbox with another upper mechbox. Currently, I am using an upgraded upper mechbox as a replacement for the previous one I had which I broke the tappet plate from. Generally, ICS M4 lower mechboxes can handle any upper mechbox you swap in, because that is one of the biggest features of the gun. Changing FPS for different environments is easy. Higher FPS for outdoor games and lower FPS for indoor games.
Personally, my new upper mechbox makes my gun shoot between 310 to 320 FPS and that is more than enough for my indoor and outdoor playing needs.
Regarding your specific question, sorry. I can't really give you an answer, because I don't know. Like I said, ask LUTNIT.